Show Me Picture of a Scorpion Biting a Baby's Legs
| Scorpions Temporal range: Early Silurian – nowadays | |
|---|---|
| | |
| Hottentotta tamulus, the Indian red scorpion | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Subphylum: | Chelicerata |
| Class: | Arachnida |
| Guild: | Scorpiones C. Fifty. Koch, 1837 |
| Families | |
| come across Taxonomy | |
| | |
| Native range of Scorpiones | |
Scorpions are predatory arachnids of the gild Scorpiones. They have viii legs, and are hands recognized past a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward bend over the back and always ending with a stinger. The evolutionary history of scorpions goes dorsum 435 meg years. They mainly alive in deserts but accept adapted to a wide range of environmental conditions, and can exist establish on all continents except Antarctica. In that location are over ii,500 described species, with 22 extant (living) families recognized to date. Their taxonomy is existence revised to account for 21st-century genomic studies.
Scorpions primarily prey on insects and other invertebrates, but some species hunt vertebrates. They apply their pincers to restrain and kill prey, or to forestall their own predation. The venomous sting is used for offense and defense force. During courting, the male and female grasp each other'due south pincers and trip the light fantastic while he tries to movement her onto his sperm package. All known species give live nascence and the female cares for the young as their exoskeletons harden, transporting them on her back. The exoskeleton contains fluorescent chemicals and glows nether ultraviolet light.
The vast majority of species practice not seriously threaten humans, and healthy adults usually do not need medical handling after a sting. Well-nigh 25 species (fewer than one percent) have venom capable of killing a human, which happens frequently in parts of the world they live, primarily where access to medical handling is unlikely.
Scorpions appear in art, sociology, mythology, and commercial brands. Scorpion motifs are woven into kilim carpets for protection from their sting. Scorpius is the name of a constellation; the corresponding astrological sign is Scorpio. A classical myth about Scorpius tells how the behemothic scorpion and its enemy Orion became constellations on opposite sides of the heaven.
Etymology
The discussion "scorpion" originated in Center English betwixt 1175 and 1225 AD from Old French scorpion ,[1] or from Italian scorpione , both derived from the Latin scorpio , equivalent to scorpius ,[2] which is the romanization of the Greek σκορπίος – skorpíos ,[three] ultimately from Proto-Indo-European root *(southward)ker- meaning "to cut", cf. "shear".[4]
Evolution
Fossil record
Scorpion fossils accept been found in many strata, including marine Silurian and estuarine Devonian deposits, coal deposits from the Carboniferous Period and in amber. Whether the early scorpions were marine or terrestrial has been debated, though they had book lungs like modern terrestrial species.[six] [seven] [8] [9] Over 100 fossil species of scorpion have been described.[10] The oldest found as of 2021 is Dolichophonus loudonensis, which lived during the Silurian, in nowadays-day Scotland.[11] Gondwanascorpio from the Devonian is amidst the primeval-known terrestrial animals on the Gondwana supercontinent.[12]
Phylogeny
The Scorpiones are a clade within the pulmonate Arachnida (those with volume lungs). Arachnida is placed within the Chelicerata, a subphylum of Arthropoda that contains sea spiders and horseshoe crabs, alongside terrestrial animals without book lungs such every bit ticks and harvestmen.[six] The extinct Eurypterida, sometimes chosen sea scorpions, though they were non all marine, are not scorpions; their grasping pincers were chelicerae, non homologous with the pincers (second appendages) of scorpions.[13] Scorpiones is sis to the Tetrapulmonata, a terrestrial group of pulmonates containing the spiders and whip scorpions. This 2019 cladogram summarizes:[6]
The internal phylogeny of the scorpions has been debated,[six] just genomic assay consistently places the Bothriuridae equally sister to a clade consisting of Scorpionoidea and Chactoidea. The scorpions diversified between the Devonian and the early Carboniferous. The main sectionalisation is into the clades Buthida and Iurida. The Bothriuridae diverged starting before temperate Gondwana broke up into separate land masses, completed by the Jurassic. The Iuroidea and Chactoidea are both seen not to exist single clades, and are shown equally "paraphyletic" (with quotation marks) in this 2018 cladogram.[14]
Taxonomy
Carl Linnaeus described six species of scorpion in his genus Scorpio in 1758 and 1767; three of these are now considered valid and are chosen Scorpio maurus, Androctonus australis, and Euscorpius carpathicus; the other three are dubious names. He placed the scorpions among his "Insecta aptera" (wingless insects), a group that included Crustacea, Arachnida and Myriapoda.[xv] In 1801, Jean-Baptiste Lamarck divided up the "Insecta aptera", creating the taxon Arachnides for spiders, scorpions, and acari (mites and ticks), though it also contained the Thysanura (thrips), Myriapoda and parasites such equally lice.[sixteen] German arachnologist Carl Ludwig Koch created the club Scorpiones in 1837. He divided information technology into iv families, the vi-eyed scorpions "Scorpionides", the eight-eyed scorpions "Buthides", the ten-eyed scorpions "Centrurides", and the twelve-eyed scorpions "Androctonides".[17]
More recently, some twenty-two families containing over 2,500 species of scorpions have been described, with many additions and much reorganization of taxa in the 21st century.[18] [half-dozen] [19] At that place are over 100 described taxa of fossil scorpions.[10] This classification is based on Soleglad and Fet (2003),[20] which replaced Stockwell'south older, unpublished classification.[21] Further taxonomic changes are from papers by Soleglad et al. (2005).[22] [23]
The extant taxa to the rank of family (numbers of species in parentheses[18]) are:
- Order Scorpiones
- Parvorder Pseudochactida Soleglad & Fet, 2003
- Superfamily Pseudochactoidea Gromov, 1998
- Family Pseudochactidae Gromov, 1998 (ane sp.) (Central Asian scorpions of semi-savanna habitats)
- Superfamily Pseudochactoidea Gromov, 1998
- Parvorder Buthida Soleglad & Fet, 2003
- Superfamily Buthoidea C. Fifty. Koch, 1837
- Family unit Buthidae C. 50. Koch, 1837 (1209 spp.) (thick-tailed scorpions, including the most dangerous species)
- Family unit Microcharmidae Lourenço, 1996, 2019 (17 spp.) (African scorpions of boiling woods leafage litter)
- Superfamily Buthoidea C. Fifty. Koch, 1837
- Parvorder Chaerilida Soleglad & Fet, 2003
- Superfamily Chaeriloidea Pocock, 1893
- Family unit Chaerilidae Pocock, 1893 (51 spp.) (South and Southeast Asian scorpions of not-arid places)
- Superfamily Chaeriloidea Pocock, 1893
- Parvorder Iurida Soleglad & Fet, 2003
- Superfamily Chactoidea Pocock, 1893
- Family Akravidae Levy, 2007 (1 sp.) (cave-dwelling house scorpions of State of israel)
- Family unit Belisariidae Lourenço, 1998 (three spp.) (cavern-related scorpions of Southern Europe)
- Family Chactidae Pocock, 1893 (209 spp.) (New World scorpions, membership under revision)
- Family Euscorpiidae Laurie, 1896 (170 spp.) (harmless scorpions of the Americas, Eurasia, and Due north Africa)
- Family Superstitioniidae Stahnke, 1940 (1 sp.) (cavern scorpions of Mexico and Southwestern Usa)
- Family Troglotayosicidae Lourenço, 1998 (4 spp.) (cave-related scorpions of Southward America)
- Family Typhlochactidae Mitchell, 1971 (eleven spp.) (cave-related scorpions of Eastern United mexican states)
- Family unit Vaejovidae Thorell, 1876 (222 spp.) (New World scorpions)
- Superfamily Iuroidea Thorell, 1876
- Family unit Caraboctonidae Kraepelin, 1905 (23 spp.) (hairy scorpions)
- Family Hadruridae Stahnke, 1974 (9 spp.) (big Due north American scorpions)
- Family Iuridae Thorell, 1876 (21 spp.) (scorpions with a big molar on inner side of moveable claw)
- Superfamily Scorpionoidea Latreille, 1802
- Family Bothriuridae Simon, 1880 (158 spp.) (Southern hemisphere tropical and temperate scorpions)
- Family Hemiscorpiidae Pocock, 1893 (16 spp.) (rock, creeping, or tree scorpions of the Middle East)
- Family Hormuridae Laurie, 1896 (92 spp.) (flattened, cleft-living scorpions of Southeast Asia and Australia)
- Family Rugodentidae Bastawade et al., 2005 (1 sp.) (burrowing scorpions of Republic of india)
- Family unit Scorpionidae Latreille, 1802 (183 spp.) (burrowing or stake-legged scorpions)
- Family Diplocentridae Karsch, 1880 (134 spp.) (closely related to and sometimes placed in Scorpionidae, but have spine on telson)
- Family Heteroscorpionidae Kraepelin, 1905 (6 spp.) (scorpions of Madagascar)
- Superfamily Chactoidea Pocock, 1893
Geographical distribution
Scorpions are plant on all continents except Antarctica. The diverseness of scorpions is greatest in subtropical areas; it decreases toward the poles and equator, though scorpions are plant in the tropics. Scorpions did not occur naturally in Britain, New Zealand and some of the islands in Oceania, only have at present been accidentally introduced into these places by humans.[24] Five colonies of Euscorpius flavicaudis have established themselves since the late 19th century in Sheerness in England at 51°N,[25] [26] [27] while Paruroctonus boreus lives as far north as Red Deer, Alberta, at 52°N.[28] A few species are on the IUCN Blood-red List; Lychas braueri is classed as critically endangered (2014), Isometrus deharvengi as endangered (2016) and Chiromachus ochropus as vulnerable (2014).[29] [30] [31]
Scorpions are xerocoles, meaning they primarily live in deserts, but they can be found in most every terrestrial habitat including high-acme mountains, caves, and intertidal zones. They are largely absent from boreal ecosystems such as the tundra, high-altitude taiga, and mountain tops.[32] [6] The highest altitude reached by a scorpion is 5,500 meters (eighteen,000 ft) in the Andes, for Orobothriurus crassimanus.[33]
As regards microhabitats, scorpions may be ground-dwelling, tree-loving, rock-loving or sand-loving. Some species, such as Vaejovis janssi, are versatile and are found in all habitats on Socorro Island, Baja California, while others such as Euscorpius carpathicus, owned to the littoral zone of rivers in Romania, occupy specialized niches.[34] [35]
Morphology
Scorpions range in size from the 8.5 mm (0.33 in) Typhlochactas mitchelli of Typhlochactidae,[34] to the 23 cm (ix.ane in) Heterometrus swammerdami of Scorpionidae.[36] The body of a scorpion is divided into two parts or tagmata: the cephalothorax or prosoma, and the abdomen or opisthosoma.[a] The opisthosoma is subdivided into a broad anterior portion, the mesosoma or pre-abdomen, and a narrow tail-similar posterior, the metasoma or mail-belly.[38] External differences between the sexes are not obvious in most species. In some, the metasoma is more elongated in males than females.[39]
Cephalothorax
The cephalothorax comprises the carapace, eyes, chelicerae (mouth parts), pedipalps (which have chelae, commonly called claws or pincers) and four pairs of walking legs. Scorpions accept two eyes on the top of the cephalothorax, and usually two to 5 pairs of optics along the front end corners of the cephalothorax. While unable to form sharp images, their central eyes are amongst the about light sensitive in the animal kingdom, particularly in dim lite, and makes it possible for nocturnal species to use starlight to navigate at night.[40] The chelicerae are at the front and underneath the carapace. They are pincer-similar and accept 3 segments and sharp "teeth".[41] [42] The brain of a scorpion is in the back of the cephalothorax, but to a higher place the esophagus.[43] As in other arachnids, the nervous arrangement is highly concentrated in the cephalothorax, but has a long ventral nerve cord with segmented ganglia which may be a archaic trait.[44]
The pedipalp is a segmented, clawed bagginess used for prey immobilization, defense force and sensory purposes. The segments of the pedipalp (from closest to the body outward) are coxa, trochanter, femur, patella, tibia (including the stock-still hook and the manus) and tarsus (moveable claw). A scorpion has darkened or granular raised linear ridges, called "keels" or "carinae" on the pedipalp segments and on other parts of the trunk; these are useful as taxonomic characters.[45] Unlike those of some other arachnids, the legs have non been modified for other purposes, though they may occasionally be used for digging, and females may apply them to take hold of emerging immature. The legs are covered in proprioceptors, bristles and sensory setae.[46] Depending on the species, the legs may have spines and spurs.[47]
Mesosoma
Ventral view: the pectines accept a comblike structure in an inverted V shape.
The mesosoma or preabdomen is the broad part of the opisthosoma.[38] It consists of the anterior seven somites (segments) of the opisthosoma, each covered dorsally by a sclerotized plate, its tergite. Ventrally, somites 3 to seven are armored with matching plates chosen sternites. The ventral side of somite one has a pair of genital opercula covering the gonopore. Sternite 2 forms the basal plate begetting the pectines,[48] which part as sensory organs.[49]
The next four somites, 3 to half-dozen, all bear pairs of spiracles. They serve every bit openings for the scorpion's respiratory organs, known every bit volume lungs. The spiracle openings may be slits, circular, elliptical or oval according to the species.[fifty] [51] There are thus 4 pairs of book lungs; each consists of some 140 to 150 thin lamellae filled with air inside a pulmonary chamber, connected on the ventral side to an atrial chamber which opens into a spiracle. Bristles hold the lamellae apart. A muscle opens the spiracle and widens the atrial chamber; dorsoventral muscles contract to compress the pulmonary bedroom, forcing air out, and relax to allow the chamber to refill.[52] The 7th and concluding somite does not comport appendages or any other meaning external structures.[fifty]
The mesosoma contains the center or "dorsal vessel" which is the center of the scorpion'due south open circulatory system. The center is continuous with a deep arterial system which spreads throughout the body. Sinuses return deoxygenated claret (hemolymph) to the centre; the blood is re-oxygenated by cardiac pores. The mesosoma also contains the reproductive organization. The female gonads are fabricated of 3 or four tubes that run parallel to each other and are connected by two to iv transverse anastomoses. These tubes are the sites for both oocyte formation and embryonic development. They connect to two oviducts which connect to a single atrium leading to the genital orifice.[53] Males have two gonads made of two cylindrical tubes with a ladder-similar configuration; they contain cysts which produce spermatozoa. Both tubes end in a spermiduct, one on each side of the mesosoma. They connect to glandular symmetrical structures chosen paraxial organs, which end at the genital orifice. These secrete chitin-based structures which come together to form the spermatophore.[54] [55]
Metasoma
The "tail" or metasoma consists of five segments and the telson, which is not strictly a segment. The v segments are merely body rings; they lack apparent sterna or terga, and go larger distally. These segments have keels, setae and bristles which may exist used for taxonomic classification. The anus is at the distal and ventral cease of the concluding segment, and is encircled past four anal papillae and the anal arch.[50] The tails of some species contain light receptors.[40]
The telson includes the vesicle, which contains a symmetrical pair of venom glands. Externally information technology bears the curved stinger, the hypodermic aculeus, equipped with sensory hairs. Each of the venom glands has its own duct to convey its secretion along the aculeus from the seedling of the gland to immediately near of the tip, where each of the paired ducts has its ain venom pore.[56] An extrinsic muscle organization in the tail moves it forward and propels and penetrates with the aculeus, while an intrinsic muscle system attached to the glands pumps venom through the stinger into the intended victim.[57] The stinger contains metalloproteins with zinc, hardening the tip.[58] The optimal stinging angle is around xxx degrees relative to the tip.[59]
Biology
Most scorpion species are nocturnal or crepuscular, finding shelter during the twenty-four hour period in burrows, cracks in rocks and tree bark.[60] Many species dig a shelter underneath stones a few centimeters long. Some may employ burrows made by other animals including spiders, reptiles and small mammals. Other species dig their own burrows which vary in complexity and depth. Hadrurus species dig burrows as over 2 k (half dozen ft vii in) deep. Digging is done using the oral cavity parts, claws and legs. In several species, particularly of the family unit Buthidae, individuals may gather in the same shelter; bawl scorpions may aggregate up to thirty individuals. In some species, families of females and young sometimes amass.[61]
Scorpions adopt areas where the temperature remains in the range of xi–40 °C (52–104 °F), but may survive temperatures from well below freezing to desert heat.[62] [63] Scorpions tin can withstand intense heat: Leiurus quinquestriatus, Scorpio maurus and Hadrurus arizonensis can live in temperatures of 45–l °C (113–122 °F) if they are sufficiently hydrated. Desert species must deal with the extreme changes in temperature from day to dark or between seasons; Pectinibuthus birulai lives in a temperature range of −30–50 °C (−22–122 °F). Scorpions that alive exterior deserts adopt lower temperatures. The ability to resist cold may be related to the increase in the sugar trehalose when the temperature drops. Some species hibernate.[64] Scorpions announced to have resistance to ionizing radiation. This was discovered in the early 1960s when scorpions were found to exist among the few animals to survive nuclear tests at Reggane, People's democratic republic of algeria.[65]
Desert scorpions have several adaptations for water conservation. They excrete insoluble compounds such equally xanthine, guanine, and uric acrid, not requiring water for their removal from the torso. Guanine is the main component and maximizes the amount of nitrogen excreted. A scorpion's cuticle holds in moisture via lipids and waxes from epidermal glands, and protects confronting ultraviolet radiations. Even when dehydrated, a scorpion can tolerate high osmotic force per unit area in its blood.[66] Desert scorpions get most of their moisture from the food they eat but some can absorb water from the boiling soil. Species that live in denser vegetation and in more moderate temperatures will drink h2o on plants and in puddles.[67]
A few scorpions squirt venom to deter predators.
A scorpion uses its stinger both for killing prey and defense. Some species make direct, quick strikes with their tails while others make slower, more than circular strikes which can more than easily return the stinger to a position where it tin can strike over again. Leiurus quinquestriatus can whip its tail at a speed of up to 128 cm/s (fifty in/s) in a defensive strike.[68]
Mortality and defense
Scorpions may be attacked by other arthropods like ants, spiders, solifugids and centipedes. Major predators include frogs, lizards, snakes, birds, and mammals.[69] Meerkats are somewhat specialized in preying on scorpions, biting off their stingers and being immune to their venom.[70] [71] Other predators adapted for hunting scorpions include the grasshopper mouse and desert long-eared bat, which are also allowed to their venom.[72] [73] In one report, 70% of the latter's droppings contained scorpion fragments.[73] Scorpions host parasites including mites, scuttle flies, nematodes and some leaner. The immune system of scorpions gives them resistance to infection by many types of bacteria.[74]
When threatened, a scorpion raises its claws and tail in a defensive posture. Some species stridulate to warn off predators past rubbing certain hairs, the stinger or the claws.[69] Certain species have a preference for using either the claws or stinger every bit defense force, depending on the size of the appendages.[75] A few scorpions, such as Parabuthus, Centruroides margaritatus, and Hadrurus arizonensis, squirt venom in a narrow jet every bit far as ane meter (3.3 ft) to warn off potential predators, possibly injuring them in the eyes.[76] Some Ananteris species can shed parts of their tail to escape predators. The parts practice not grow back, leaving them unable to sting and defecate, but they tin still take hold of small prey and reproduce for at least eight months afterward.[77]
Diet and feeding
Scorpions generally prey on insects, particularly grasshoppers, crickets, termites, beetles and wasps. Other casualty include spiders, solifugids, woodlice and even small-scale vertebrates including lizards, snakes and mammals. Species with big claws may prey on earthworms and mollusks. The majority of species are opportunistic and eat a diverseness of casualty though some may exist highly specialized; Isometroides vescus specializes on burrowing spiders. Prey size depends on the size of the species. Several scorpion species are sit-and-wait predators, which involves them waiting for prey at or near the archway to their couch. Others actively seek them out. Scorpions detect their prey with mechanoreceptive and chemoreceptive hairs on their bodies and capture them with their claws. Minor animals are merely killed with the claws, particularly by large-clawed species. Larger and more than aggressive casualty is given a sting.[78] [79]
Scorpions, like other arachnids, digest their food externally. The chelicerae, which are very sharp, are used to pull small amounts of nutrient off the casualty particular into a pre-rima oris beneath the chelicerae and carapace. The digestive juices from the gut are egested onto the nutrient, and the digested food is then sucked into the gut in liquid grade. Any solid indigestible matter (such every bit exoskeleton fragments) is trapped by setae in the pre-oral cavity and ejected. The sucked-in nutrient is pumped into the midgut by the throat, where information technology is further digested. The waste passes through the hindgut and out of the anus. Scorpions tin consume large amounts of food during ane repast. They have an efficient food storage organ and a very low metabolic rate, and a relatively inactive lifestyle. This enables some to survive half dozen to twelve months of starvation.[80]
Mating
Male and female person scorpion during promenade à deux
Virtually scorpions reproduce sexually, with male and female person individuals; species in some genera, such equally Hottentotta and Tityus, and the species Centruroides gracilis, Liocheles australasiae, and Ananteris coineaui have been reported, not necessarily reliably, to reproduce through parthenogenesis, in which unfertilized eggs develop into living embryos.[81] Receptive females produce pheromones which are picked upwardly by wandering males using their pectines to comb the substrate. Males brainstorm courting past moving their bodies back and forth, without moving the legs, a behavior known equally juddering. This appears to produce ground vibrations that are picked up past the female.[54]
The pair then make contact using their pedipalps, and perform a dance called the promenade à deux (French for "a walk for 2"). In this dance, the male and female move back and forth while facing each other, every bit the male searches for a suitable identify to deposit his spermatophore. The courting ritual tin can involve several other behaviors such equally a cheliceral kiss, in which the male person and female grasp each other'southward mouth-parts, arbre droit ("upright tree") where the partners elevate their posteriors and rub their tails together, and sexual stinging, in which the male stings the female person in the chelae or mesosoma to subdue her. The dance tin last from a few minutes to several hours.[82] [83]
When the male has located a suitably stable substrate, such as hard ground, agglomerated sand, rock, or tree bark, he deposits the spermatophore and guides the female person over it. This allows the spermatophore to enter her genital opercula, which triggers release of the sperm, thus fertilizing the female person. A mating plug then forms in the female to forbid her from mating again before the young are built-in. The male person and female then abruptly split up.[84] [85] Sexual cannibalism after mating has only been reported anecdotally in scorpions.[86]
Nativity and development
Gestation in scorpions can terminal for over a year in some species.[87] They accept two types of embryonic development; apoikogenic and katoikogenic. In the apoikogenic arrangement, which is mainly establish in the Buthidae, embryos develop in yolk-rich eggs inside follicles. The katoikogenic system is documented in Hemiscorpiidae, Scorpionidae and Diplocentridae, and involves the embryos developing in a diverticulum which has a teat-like structure for them to feed though.[88] Unlike the majority of arachnids, which are oviparous, hatching from eggs, scorpions seem to be universally viviparous, with alive births.[89] They are unusual among terrestrial arthropods in the amount of intendance a female gives to her offspring.[ninety] The size of a brood varies by species, from iii to over 100.[91] The body size of scorpions is not correlated either with breed size or with life bike length.[92]
Before giving birth, the female elevates the forepart of her body and positions her pedipalps and front legs nether her to grab the immature ("birth basket"). The immature emerge one by one from the genital opercula, expel the embryonic membrane, if any, and are placed on the mother'southward back where they remain until they have gone through at least ane molt. The period before the showtime molt is called the pro-juvenile phase; the young are unable to feed or sting, but have suckers on their tarsi, used to hold on to their mother. This catamenia lasts 5 to 25 days, depending on the species. The brood molt for the first time simultaneously in a process that lasts six to 8 hours, marking the kickoff of the juvenile phase.[91]
Juvenile stages or instars mostly resemble smaller versions of adults, with fully adult pincers, hairs and stingers. They are withal soft and lack pigments, and thus continue to ride on their mother's back for protection. They become harder and more pigmented over the next couple of days. They may leave their female parent temporarily, returning when they sense potential danger. In one case the exoskeleton is fully hardened, the young tin can hunt prey on their ain and may before long go out their mother.[93] A scorpion may molt six times on average earlier reaching maturity, which may not occur until it is 6 to 83 months old, depending on the species. Some species may live up to 25 years.[87]
Fluorescence
The female parent glows bright teal, the babies a ho-hum grey.
Scorpions glow a vibrant blue-greenish when exposed to certain wavelengths ranges of ultraviolet low-cal such as that produced past a black light, due to fluorescent chemicals such equally beta-carboline in the cuticle. Appropriately, a paw-held ultraviolet lamp has long been a standard tool for nocturnal field surveys of these animals. Fluorescence occurs as a result of sclerotization and increases in intensity with each successive instar.[94] This fluorescence may have an active function in the scorpion'south power to notice light.[95]
Human relationship with humans
Stings
Scorpion venom serves to kill or paralyze prey rapidly. The stings of many species are uncomfortable, but just 25 species have venom that is deadly to humans. Those species belong to the family Buthidae, including Leiurus quinquestriatus, Hottentotta spp., Centruroides spp., and Androctonus spp.[34] People with allergies are especially at adventure;[96] otherwise, outset aid is symptomatic, with analgesia. Cases of very high claret force per unit area are treated with medications that salve anxiety and relax the blood vessels.[97] [98] Scorpion envenomation with high morbidity and mortality is usually due to either excessive autonomic activity and cardiovascular toxic effects, or neuromuscular toxic effects. Antivenom is the specific handling for scorpion envenomation combined with supportive measures including vasodilators in patients with cardiovascular toxic effects, and benzodiazepines when at that place is neuromuscular involvement. Although rare, severe hypersensitivity reactions including anaphylaxis to scorpion antivenin are possible.[99]
Scorpion stings are a public health problem, particularly in the tropical and subtropical regions of the Americas, North Africa, the Middle Due east and India. Effectually 1.v million scorpion envenomations occur each year with around 2,600 deaths.[100] [101] [102] Mexico is one of the most affected countries, with the highest biodiversity of scorpions in the world, some 200,000 envenomations per year and at least 300 deaths.[103] [104]
Efforts are fabricated to prevent envenomation and to control scorpion populations. Prevention encompasses personal activities such as checking shoes and clothes before putting them on, non walking in blank anxiety or sandals, and filling in holes and cracks where scorpions might nest. Street lighting reduces scorpion activity. Command may involve the apply of insecticides such as pyrethroids, or gathering scorpions manually with the help of ultraviolet lights. Domestic predators of scorpions, such as chickens and turkeys, can help to reduce the run a risk to a household.[100] [101]
Potential medicinal use
Scorpion venom is a mixture of neurotoxins; well-nigh of these are peptides, chains of amino acids.[106] Many of them interfere with membrane channels that transport sodium, potassium, calcium, or chloride ions. These channels are essential for nerve conduction, muscle contraction and many other biological processes. Some of these molecules may be useful in medical inquiry and might lead to the evolution of new illness treatments. Among their potential therapeutic uses are as analgesic, anti-cancer, antibacterial, antifungal, antiviral, antiparasitic, bradykinin-potentiating, and immunosuppressive drugs. Every bit of 2020, no scorpion toxin-based drug is on sale, though chlorotoxin is being trialled for use against glioma, a brain cancer.[105]
Consumption
Scorpions are eaten by people in West Africa, Myanmar[107] and East asia. Fried scorpion is traditionally eaten in Shandong, Prc.[108] There, scorpions can be cooked and eaten in a variety of ways, including roasting, frying, grilling, raw, or alive. The stingers are typically non removed, since straight and sustained heat negates the harmful effects of the venom.[109] In Thailand, scorpions are not eaten as often as other arthropods, such as grasshoppers, only they are sometimes fried as street food.[110] They are used in Vietnam to make snake vino (scorpion wine).[111]
Pets
Scorpions are often kept as pets. They are relatively unproblematic to go on, the principal requirements beingness a secure enclosure such every bit a glass terrarium with a lockable lid and the appropriate temperature and humidity for the called species, which typically means installing a heating mat and spraying regularly with a little water. The substrate needs to resemble that of the species' natural environment, such every bit peat for forest species, or lateritic sand for burrowing desert species. Scorpions in the genera Pandinus and Heterometrus are docile enough to handle. A large Pandinus may swallow up to three crickets each week. Cannibalism is more common in captivity than in the wild and tin can be minimized by providing many small shelters within the enclosure and ensuring there is plenty of prey.[112] [113] The pet trade has threatened wild populations of some scorpion species, specially Androctonus australis and Pandinus imperator.[114]
Civilization
-
Late period bronze figure of Isis-Serket
-
A scorpion motif (2 types shown) was often woven into Turkish kilim flatweave carpets, for protection from their sting.[115]
The scorpion is a culturally pregnant brute, appearing as a motif in art, particularly in Islamic art in the Middle East.[116] A scorpion motif is frequently woven into Turkish kilim flatweave carpets, for protection from their sting.[115] The scorpion is perceived both as an embodiment of evil and equally a protective strength such as a dervish's powers to combat evil.[116] In Muslim folklore, the scorpion portrays homo sexuality.[116] Scorpions are used in folk medicine in Southern asia, peculiarly in antidotes for scorpion stings.[116]
1 of the earliest occurrences of the scorpion in culture is its inclusion, as Scorpio, in the 12 signs of the Zodiac by Babylonian astronomers during the Chaldean menstruation. This was then taken up by western star divination; in astronomy the corresponding constellation is named Scorpius.[117] In ancient Egypt, the goddess Serket, who protected the Pharaoh, was ofttimes depicted as a scorpion.[118] In ancient Greece, a warrior's shield sometimes carried a scorpion device, as seen in red-figure pottery from the 5th century BC.[119] In Greek mythology, Artemis or Gaia sent a giant scorpion to kill the hunter Orion, who had said he would kill all the earth's animals. Orion and the scorpion both became constellations; every bit enemies they were placed on reverse sides of the world, and then when one rises in the sky, the other sets.[120] [121] Scorpions are mentioned in the Bible and the Talmud as symbols of danger and maliciousness.[121]
The legend of The Scorpion and the Frog has been interpreted as showing that vicious people cannot resist pain others, fifty-fifty when it is not in their interests.[122] More recently, the activity in John Steinbeck'south 1947 novella The Pearl centers on a poor pearl fisherman'due south attempts to save his infant son from a scorpion sting, but to lose him to human violence.[123] Scorpions take equally appeared in western artforms including motion picture and poesy: the surrealist filmmaker Luis Buñuel made symbolic utilize of scorpions in his 1930 classic L'Age d'or (The Aureate Historic period),[124] while Stevie Smith's last collection of poems was entitled Scorpion and other Poems.[125] A diversity of martial arts films and video games have been entitled Scorpion Rex.[126] [127] [128]
Scorpion pose in yoga has 1 or both legs pointing forward over the head, like a scorpion's tail.[129]
Since classical times, the scorpion with its powerful stinger has been used to provide a name for weapons. In the Roman army, the scorpio was a torsion siege engine used to shoot a projectile.[130] The British Army's FV101 Scorpion was an armored reconnaissance vehicle or calorie-free tank in service from 1972 to 1994.[131] A version of the Matilda II tank, fitted with a flail to clear mines, was named the Matilda Scorpion.[132] Several ships of the Royal Navy and of the United states of america Navy accept been named Scorpion including an 18-gun sloop in 1803,[133] a turret transport in 1863,[134] a patrol yacht in 1898,[135] a destroyer in 1910,[136] and a nuclear submarine in 1960.[137]
The scorpion has served as the name or symbol of products and brands including Italian republic's Abarth racing cars[138] and a Montesa scrambler motorcycle.[139] A mitt- or forearm-balancing asana in modern yoga as do with the back arched and 1 or both legs pointing forrad over the caput in the manner of the scorpion's tail is chosen Scorpion pose.[140] [129]
Notes
- ^ Equally there is currently neither paleontological nor embryological evidence that arachnids ever had a split thorax-like partitioning, in that location exists an statement against the validity of the term cephalothorax, which ways fused cephalon (head) and the thorax. Similarly, arguments can be formed against use of the term abdomen, as the opisthosoma of all scorpions contains a center and book lungs, organs atypical of an abdomen.[37]
References
- ^ "Scorpion". The American Heritage Dictionary of the English Language, 5th Edition. 2016. Archived from the original on 22 November 2020. Retrieved vii Dec 2020.
- ^ "Scorpion". Dictionary.com. Archived from the original on fifteen Nov 2015. Retrieved 7 December 2020.
- ^ σκορπίος . Liddell, Henry George; Scott, Robert; A Greek–English Dictionary at the Perseus Project.
- ^ Harper, Douglas. "scorpion". Online Etymology Dictionary.
- ^ Pocock, R. I. (1901). . The Quarterly Journal of Microscopical Science. 44. Plate nineteen – via Wikisource.
- ^ a b c d e f Howard, Richard J.; Edgecombe, Gregory D.; Legg, David A.; Pisani, Davide; Lozano-Fernandez, Jesus (2019). "Exploring the Evolution and Terrestrialization of Scorpions (Arachnida: Scorpiones) with Rocks and Clocks". Organisms Diversity & Evolution. 19 (1): 71–86. doi:x.1007/s13127-019-00390-7. ISSN 1439-6092.
- ^ Scholtz, Gerhard; Kamenz, Carsten (2006). "The Volume Lungs of Scorpiones and Tetrapulmonata (Chelicerata, Arachnida): Evidence for Homology and a Unmarried Terrestrialisation Consequence of a Mutual Arachnid Ancestor". Zoology. 109 (1): 2–13. doi:10.1016/j.zool.2005.06.003. PMID 16386884.
- ^ Dunlop, Jason A.; Tetlie, O. Erik; Prendini, Lorenzo (2008). "Reinterpretation of the Silurian Scorpion Proscorpius osborni (Whitfield): Integrating Data from Palaeozoic and Contempo Scorpions". Palaeontology. 51 (2): 303–320. doi:10.1111/j.1475-4983.2007.00749.x.
- ^ Kühl, Grand.; Bergmann, A.; Dunlop, J.; Garwood, R. J.; Rust, J. (2012). "Redescription and Palaeobiology of Palaeoscorpius devonicus Lehmann, 1944 from the Lower Devonian Hunsrück Slate of Frg". Palaeontology. 55 (4): 775–787. doi:10.1111/j.1475-4983.2012.01152.10.
- ^ a b Dunlop, J. A.; Penney, D. (2012). Fossil Arachnids. Siri Scientific Printing. p. 23. ISBN978-0956779540.
- ^ Anderson, Evan P.; Schiffbauer, James D.; Jacquet, Sarah M.; Lamsdell, James C.; Kluessendorf, Joanne; Mikulic, Donald G. (2021). "Stranger than a scorpion: a reassessment of Parioscorpio venator, a problematic arthropod from the Llandoverian Waukesha Lagerstätte". Palaeontology. 64 (3): 429–474. doi:10.1111/pala.12534. ISSN 1475-4983. S2CID 234812878. Archived from the original on 20 April 2021. Retrieved twenty Apr 2021.
- ^ Gess, R. Westward. (2013). "The Earliest Record of Terrestrial Animals in Gondwana: a Scorpion from the Famennian (Belatedly Devonian) Witpoort Formation of S Africa". African Invertebrates. 54 (two): 373–379. doi:ten.5733/afin.054.0206. Archived from the original on 6 September 2013. Retrieved 28 Baronial 2013.
- ^ Waggoner, B. M. (12 October 1999). "Eurypterida: Morphology". Academy of California Museum of Paleontology Berkeley. Archived from the original on 23 Oct 2020. Retrieved 20 Oct 2020.
- ^ Sharma, Prashant P.; Baker, Caitlin One thousand.; Cosgrove, Julia G.; Johnson, Joanne E.; Oberski, Jill T.; Raven, Robert J.; Harvey, Marker S.; Boyer, Sarah L.; Giribet, Gonzalo (2018). "A Revised Dated Phylogeny of Scorpions: Phylogenomic Back up for Ancient Difference of the Temperate Gondwanan Family Bothriuridae". Molecular Phylogenetics and Evolution. 122: 37–45. doi:10.1016/j.ympev.2018.01.003. ISSN 1055-7903. PMID 29366829.
- ^ Fet, 5.; Braunwalder, Thou. Due east.; Cameron, H. D. (2002). "Scorpions (Arachnida, Scorpiones) Described by Linnaeus" (PDF). Bulletin of the British Arachnological Society. 12 (4): 176–182. Archived (PDF) from the original on 26 September 2020. Retrieved 20 October 2020.
- ^ Burmeister, Carl Hermann C.; Shuckard, W. East. (trans) (1836). A Transmission of Entomology. pp. 613ff. Archived from the original on 31 May 2021. Retrieved 20 Oct 2020.
- ^ Koch, Carl Ludwig (1837). Übersicht des Arachnidensystems (in German). C. H. Zeh. pp. 86–92.
- ^ a b "The Scorpion Files". Jan Ove Rein. Archived from the original on iii October 2020. Retrieved 15 Baronial 2020.
- ^ Kovařík, František (2009). "Illustrated Catalog of Scorpions, Role I" (PDF). Archived (PDF) from the original on 26 October 2010. Retrieved 22 January 2011.
- ^ Soleglad, Michael E.; Fet, Victor (2003). "High-level Systematics and Phylogeny of the Extant Scorpions (Scorpiones: Orthosterni)" (multiple parts). Euscorpius. 11: 1–175. Archived from the original on 8 December 2019. Retrieved 13 June 2008.
- ^ Stockwell, Scott A. (1989). Revision of the Phylogeny and Higher Classification of Scorpions (Chelicerata) (PhD thesis). University of California, Berkeley.
- ^ Soleglad, Michael East.; Fet, Victor; Kovařík, F. (2005). "The Systematic Position of the Scorpion Genera Heteroscorpion Birula, 1903 and Urodacus Peters, 1861 (Scorpiones: Scorpionoidea)" (PDF). Euscorpius. 20: ane–38. Archived (PDF) from the original on 16 December 2008. Retrieved 13 June 2008.
- ^ Fet, Victor; Soleglad, Michael Eastward. (2005). "Contributions to Scorpion Systematics. I. On Recent Changes in High-level Taxonomy" (PDF). Euscorpius (31): ane–13. ISSN 1536-9307. Archived (PDF) from the original on 26 April 2018. Retrieved vii Apr 2010.
- ^ Polis 1990, p. 249.
- ^ Benton, T. G. (1992). "The Ecology of the Scorpion Euscorpius flavicaudis in England". Periodical of Zoology. 226 (3): 351–368. doi:x.1111/j.1469-7998.1992.tb07484.x.
- ^ Benton, T. G. (1991). "The Life History of Euscorpius flavicaudis (Scorpiones, Chactidae)" (PDF). The Journal of Arachnology. 19 (2): 105–110. JSTOR 3705658. Archived (PDF) from the original on 28 February 2008. Retrieved 16 January 2008.
- ^ Rein, Jan Ove (2000). "Euscorpius flavicaudis". The Scorpion Files. Norwegian University of Science and Technology. Archived from the original on 26 Apr 2018. Retrieved 13 June 2008.
- ^ Johnson, D. L. (2004). "The Northern Scorpion, Paruroctonus boreus, in Southern Alberta, 1983–2003". Arthropods of Canadian Grasslands x (PDF). Biological Survey of Canada. Archived from the original (PDF) on 29 November 2015.
- ^ Gerlach, J. (2014). "Lychas braueri". IUCN Red List of Threatened Species. 2014 . Retrieved x Dec 2020.
- ^ Deharveng, Fifty.; Bedos, A. (2016). "Isometrus deharvengi". IUCN Scarlet List of Threatened Species. 2016 . Retrieved 10 Dec 2020.
- ^ Gerlach, J. (2014). "Chiromachus ochropus". IUCN Reddish Listing of Threatened Species. 2014 . Retrieved ten December 2020.
- ^ Polis 1990, pp. 251–253.
- ^ Stockmann & Ythier 2010, p. 151.
- ^ a b c Ramel, Gordon. "The Earthlife Web: The Scorpions". The Earthlife Web. Archived from the original on 9 October 2019. Retrieved 8 Apr 2010.
- ^ Gherghel, I.; Sotek, A.; Papes, M.; Strugariu, A.; Fusu, L. (2016). "Ecology and Biogeography of the Endemic Scorpion Euscorpius carpathicus (Scorpiones: Euscorpiidae): a Multiscale Assay". Journal of Arachnology. 44 (1): 88–91. doi:10.1636/P14-22.1. S2CID 87325752.
- ^ Rubio, Manny (2000). "Commonly Bachelor Scorpions". Scorpions: Everything Virtually Purchase, Intendance, Feeding, and Housing. Barron's. pp. 26–27. ISBN978-0-7641-1224-nine.
The Guinness Volume of Records claims [...] Heterometrus swammerdami, to be the largest scorpion in the world [ix inches (23 cm)]
- ^ Shultz, Stanley; Shultz, Marguerite (2009). The Tarantula Keeper's Guide. Barron's. p. 23. ISBN978-0-7641-3885-0.
- ^ a b Polis 1990, pp. 10–xi.
- ^ Stockmann & Ythier 2010, p. 76.
- ^ a b Chakravarthy, Akshay Kumar; Sridhara, Shakunthala (2016). Arthropod Diversity and Conservation in the Tropics and Sub-tropics. Springer. p. 60. ISBN978-981-10-1518-2. Archived from the original on 31 May 2021. Retrieved 11 September 2017.
- ^ Polis 1990, pp. sixteen–17.
- ^ Stockmann 2015, p. 26.
- ^ Polis 1990, p. 38.
- ^ Polis 1990, p. 342.
- ^ Polis 1990, p. 12.
- ^ Polis 1990, p. 20.
- ^ Polis 1990, p. 74.
- ^ Polis 1990, pp. 13–14.
- ^ Knowlton, Elizabeth D.; Gaffin, Douglas D. (2011). "Functionally Redundant Peg Sensilla on the Scorpion Pecten". Journal of Comparative Physiology A. Springer. 197 (9): 895–902. doi:10.1007/s00359-011-0650-nine. ISSN 0340-7594. PMID 21647695. S2CID 22123929.
- ^ a b c Polis 1990, p. xv.
- ^ Wanninger, Andreas (2015). Evolutionary Developmental Biology of Invertebrates 3: Ecdysozoa I: Non-Tetraconata. Springer. p. 105. ISBN978-3-7091-1865-8. Archived from the original on 5 Baronial 2020. Retrieved 11 September 2017.
- ^ Polis 1990, pp. 42–44.
- ^ Stockmann 2015, pp. 45–46.
- ^ a b Stockmann 2015, p. 47.
- ^ Lautié, N.; Soranzo, L.; Lajarille, One thousand.-East.; Stockmann, R. (2007). "Paraxial Organ of a Scorpion: Structural and Ultrastructural Studies of Euscorpius tergestinus Paraxial Organ (Scorpiones, Euscorpiidae)". Invertebrate Reproduction & Development. 51 (2): 77–90. doi:10.1080/07924259.2008.9652258. S2CID 84763256.
- ^ Yigit, N.; Benli, M. (2010). "Fine Structural Analysis of the Stinger in Venom Apparatus of the Scorpion Euscorpius mingrelicus (Scorpiones: Euscorpiidae)". Periodical of Venomous Animals and Toxins Including Tropical Diseases. 16 (1): 76–86. doi:10.1590/s1678-91992010005000003.
- ^ Stockmann 2015, p. 30.
- ^ Schofield, R. M. S. (2001). "Metals in cuticular structures". In Brownell, P. H.; Polis, Thousand. A. (eds.). Scorpion Biology and Research. New York: Oxford University Press. pp. 234–256. ISBN978-0195084344.
- ^ van der Meijden, Arie; Kleinteich, Thomas (Apr 2017). "A biomechanical view on stinger multifariousness in scorpions". Periodical of Anatomy. 230 (4): 497–509. doi:x.1111/joa.12582. PMC5345679. PMID 28028798.
- ^ Stockmann 2015, pp. 40–41.
- ^ Stockmann & Ythier 2010, pp. 146, 153–154.
- ^ Hadley, Neil F. (1970). "Water Relations of the Desert Scorpion, Hadrurus arizonensis" (PDF). Periodical of Experimental Biology. 53 (3): 547–558. doi:x.1242/jeb.53.iii.547. PMID 5487163. Archived (PDF) from the original on 6 April 2008. Retrieved 30 Dec 2006.
- ^ Hoshino, K.; Moura, A. T. V.; De Paula, H. M. 1000. (2006). "Selection of Ecology Temperature by the Yellowish Scorpion Tityus serrulatus Lutz & Mello, 1922 (Scorpiones, Buthidae)". Journal of Venomous Animals and Toxins Including Tropical Diseases. 12 (one): 59–66. doi:10.1590/S1678-91992006000100005.
- ^ Stockmann 2015, pp. 42–43.
- ^ Stockmann & Ythier 2010, p. 157.
- ^ Cowles, Jillian (2018). Amazing Arachnids. Princeton University Press. p. 33. ISBN978-0-691-17658-1.
- ^ Stockmann & Ythier 2010, p. 156.
- ^ Coelho, P.; Kaliontzopoulou, A.; Rasko, Thousand.; van der Meijden, A. (2017). "A 'Striking' Human relationship: Scorpion Defensive Behaviour and its Relation to Morphology and Performance". Functional Environmental. 31 (vii): 1390–1404. doi:10.1111/1365-2435.12855.
- ^ a b Stockmann 2015, pp. 36–37.
- ^ Van Staaden, K. J. (1994). "Suricata suricatta" (PDF). Mammalian Species (483): one–8. doi:10.2307/3504085. JSTOR 3504085. Archived from the original (PDF) on fifteen March 2016.
- ^ Thornton, A.; McAuliffe, M. (2006). "Teaching in wild meerkats" (PDF). Science. 313 (5784): 227–229. Bibcode:2006Sci...313..227T. doi:x.1126/scientific discipline.1128727. PMID 16840701. S2CID 11490465. Archived from the original (PDF) on 26 February 2020.
- ^ Thompson, Benjamin (June 2018). "The Grasshopper Mouse and Bark Scorpion: Evolutionary Biological science Meets Hurting Modulation and Selective Receptor Inactivation". The Journal of Undergraduate Neuroscience Education. 16 (ii): R51–R53. PMC6057761. PMID 30057511.
- ^ a b Holderied, Chiliad.; Korine, C.; Moritz, T. (2010). "Hemprich'south Long-eared Bat (Otonycteris hemprichii) as a Predator of Scorpions: Whispering Echolocation, Passive Gleaning and Prey Selection". Journal of Comparative Physiology A. 197 (5): 425–433. doi:ten.1007/s00359-010-0608-iii. PMID 21086132. S2CID 25692517.
- ^ Stockmann 2015, pp. 38, 45.
- ^ van der Meijden, A.; Coelho, P. L.; Sousa, P.; Herrel, A. (2013). "Choose your Weapon: Defensive Behavior is Associated with Morphology and Functioning in Scorpions". PLOS ONE. 8 (eleven): e78955. Bibcode:2013PLoSO...878955V. doi:10.1371/journal.pone.0078955. PMC3827323. PMID 24236075.
- ^ Stockmann & Ythier 2010, p. 90.
- ^ Mattoni, C. I.; García-Hernández, S.; Botero-Trujillo, R.; Ochoa, J. A.; Ojanguren-Affilastro, A. A.; Pinto-da-Rocha, R.; Prendini, L. (2015). "Scorpion Sheds 'Tail' to Escape: Consequences and Implications of Autotomy in Scorpions (Buthidae: Ananteris)". PLOS I. ten (ane): e0116639. Bibcode:2015PLoSO..1016639M. doi:x.1371/journal.pone.0116639. PMC4309614. PMID 25629529. S2CID 17870490.
- ^ Stockmann 2015, pp. 35–38.
- ^ Murray, Melissa (3 December 2020). "Scorpions". Australian Museum. Archived from the original on 8 Nov 2020. Retrieved 13 December 2020.
- ^ Polis 1990, pp. 296–298.
- ^ Lourenço, Wilson R. (2008). "Parthenogenesis in Scorpions: Some History – New Data". Journal of Venomous Animals and Toxins Including Tropical Diseases. 14 (one). doi:10.1590/S1678-91992008000100003. ISSN 1678-9199.
- ^ Stockmann 2015, pp. 47–fifty.
- ^ Stockmann & Ythier 2010, pp. 126–128.
- ^ Stockmann 2015, pp. 49–50.
- ^ Stockmann & Ythier 2010, p. 129.
- ^ Peretti, A. (1999). "Sexual Cannibalism in Scorpions: Fact or Fiction?". Biological Journal of the Linnean Society. 68 (four): 485–496. doi:10.1111/j.1095-8312.1999.tb01184.x. ISSN 0024-4066.
- ^ a b Polis 1990, p. 161.
- ^ Warburg, G. R. (2010). "Reproductive System of Female Scorpion: A Fractional Review". The Anatomical Record. 293 (x): 1738–1754. doi:ten.1002/ar.21219. PMID 20687160. S2CID 25391120.
- ^ Warburg, Michael R. (5 April 2012). "Pre- and Postal service-parturial Aspects of Scorpion Reproduction: a Review". European Periodical of Entomology. 109 (two): 139–146. doi:x.14411/eje.2012.018.
- ^ Polis 1990, p. vi.
- ^ a b Lourenço, Wilson R. (2000). "Reproduction in Scorpions, with Special Reference to Parthenogenesis" (PDF). In Toft, S.; Scharff, N. (eds.). European Arachnology. Aarhus Academy Printing. pp. 74–76. ISBN978-877934-0015. Archived (PDF) from the original on 31 May 2021. Retrieved 14 Dec 2020.
- ^ Monge-Nájera, J. (2019). "Scorpion Body Size, Litter Characteristics, and Duration of the Life Cycle (Scorpiones)". Cuadernos de Investigación UNED. 11 (2): 101–104.
- ^ Stockmann 2015, p. 54.
- ^ Stachel, Shawn J.; Stockwell, Scott A.; Van Vranken, David L. (August 1999). "The Fluorescence of Scorpions and Cataractogenesis". Chemical science & Biology. half dozen (viii): 531–539. doi:10.1016/S1074-5521(99)80085-4. PMID 10421760.
- ^ Gaffinr, Douglas D.; Bumm, Lloyd A.; Taylor, Matthew S.; Popokina, Nataliya V.; Mann, Shivani (2012). "Scorpion Fluorescence and Reaction to Light". Animal Behaviour. 83 (2): 429–436. doi:x.1016/j.anbehav.2011.eleven.014. S2CID 17041988. Archived from the original on 6 November 2020. Retrieved 18 May 2018.
- ^ "Insects and Scorpions". NIOSH. 1 July 2016. Archived from the original on 29 July 2016. Retrieved 15 July 2016.
- ^ Buma, Adriaan Hopperus; Burris, David Chiliad.; Hawley, Alan; Ryan, James G.; Mahoney, Peter F. (2009). "Scorpion sting". Conflict and Catastrophe Medicine: A Practical Guide (2nd ed.). Springer. p. 518. ISBN978-1-84800-351-4. Archived from the original on 25 Nov 2016. Retrieved 5 March 2016.
- ^ "Diseases and Conditions – Scorpion Stings". Mayo Clinic. Archived from the original on 4 October 2016. Retrieved 3 July 2015.
- ^ Bhoite, R. R.; Bhoite, Thousand. R.; Bagdure, D. N.; Bawaskar, H. S. (2015). "Anaphylaxis to Scorpion Antivenin and its Direction Post-obit Envenomation past Indian Cherry Scorpion, Mesobuthus tamulus". Indian Journal of Critical Care Medicine. nineteen (9): 547–549. doi:10.4103/0972-5229.164807. PMC4578200. PMID 26430342.
- ^ a b Feola, A.; Perrone, M. A.; Piscopo, A.; Casella, F.; Della Pietra, B.; Di Mizio, G. (2020). "Dissection Findings in Case of Fatal Scorpion Sting: A Systematic Review of the Literature". Healthcare. 8 (three): 325. doi:10.3390/healthcare8030325. PMC7551928. PMID 32899951.
- ^ a b Stockmann & Ythier 2010, pp. 163–164.
- ^ Santos, Maria South.V.; Silva, Cláudio G.L.; Neto, Basílio Silva; Grangeiro Júnior, Cícero R.P.; Lopes, Victor H.G.; Teixeira Júnior, Antônio G.; Bezerra, Deryk A.; Luna, João V.C.P.; Cordeiro, Josué B.; Júnior, Jucier Gonçalves; Lima, Marcos A.P. (2016). "Clinical and Epidemiological Aspects of Scorpionism in the World: A Systematic Review". Wilderness & Environmental Medicine. 27 (4): 504–518. doi:10.1016/j.wem.2016.08.003. ISSN 1080-6032. PMID 27912864. Archived from the original on 31 May 2021. Retrieved eight December 2020.
- ^ Dehesa-Dávila, Manuel; Possani, Lourival D. (1994). "Scorpionism and Serotherapy in Mexico". Toxicon. 32 (9): 1015–1018. doi:ten.1016/0041-0101(94)90383-ii. ISSN 0041-0101. PMID 7801335.
- ^ Santibáñez-López, Carlos; Francke, Oscar; Ureta, Carolina; Possani, Lourival (2015). "Scorpions from Mexico: From Species Diversity to Venom Complication". Toxins. 8 (one): 2. doi:10.3390/toxins8010002. ISSN 2072-6651. PMC4728524. PMID 26712787.
- ^ a b Ahmadi, Shirin; Knerr, Julius M.; Argemi, Lídia; Bordon, Karla C. F.; Pucca, Manuela B.; Cerni, Felipe A.; Arantes, Eliane C.; Çalışkan, Figen; Laustsen, Andreas H. (12 May 2020). "Scorpion Venom: Detriments and Benefits". Biomedicines. MDPI AG. viii (5): 118. doi:10.3390/biomedicines8050118. ISSN 2227-9059. PMC7277529. PMID 32408604.
- ^ Rodríguez de la Vega, Ricardo C.; Vidal, Nicolas; Possani, Lourival D. (2013). "Scorpion Peptides". In Kastin, Abba J. (ed.). Handbook of Biologically Active Peptides (2nd ed.). Academic Printing. pp. 423–429. doi:10.1016/B978-0-12-385095-9.00059-ii. ISBN978-0-12-385095-9.
- ^ Stockmann & Ythier 2010, p. 147.
- ^ Forney, Matthew (11 June 2008). "Scorpions for Breakfast and Snails for Dinner". The New York Times. Archived from the original on 27 November 2020. Retrieved 23 February 2017.
- ^ Albers, Susan (fifteen May 2014). "How to Mindfully Eat a Scorpion". HuffPost. Archived from the original on 13 August 2020. Retrieved 29 July 2020.
- ^ Fernquest, Jon (30 March 2016). "Fried Scorpion Anyone?". Bangkok Post. Archived from the original on 31 May 2021. Retrieved seven December 2020.
- ^ Lachenmeier, Dirk Due west.; Anh, Pham Thi Hoang; Popova, Svetlana; Rehm, Jürgen (11 August 2009). "The Quality of Booze Products in Vietnam and Its Implications for Public Health". International Periodical of Environmental Research and Public Health. six (eight): 2090–2101. doi:10.3390/ijerph6082090. PMC2738875. PMID 19742208.
- ^ "Scorpion Caresheet". Apprentice Entomologists' Guild. Archived from the original on one Nov 2020. Retrieved 19 November 2020.
- ^ Stockmann & Ythier 2010, pp. 144, 173–177.
- ^ Pryke, Fifty. M. (2016). Scorpion. Reaktion Books. pp. 187–189. ISBN978-1780236254.
- ^ a b Erbek, Güran (1998). Kilim Catalogue No. one (1st ed.). May Selçuk A. S. "Motifs", before page i.
- ^ a b c d Frembgen, Jürgen Wasim (2004). "The Scorpion in Muslim Folklore". Asian Folklore Studies. 63 (one): 95–123. Archived from the original on 16 March 2014. Retrieved 12 August 2014.
- ^ Polis 1990, p. 462.
- ^ "Serqet". British Museum. Archived from the original on one December 2020. Retrieved x Dec 2020.
- ^ attrib. to "The Achilles Painter". "Pelike". British Museum. Archived from the original on 3 October 2020. Retrieved 24 August 2020.
in his left [hand] he carries a long spear and a shield with device in black silhouette of a scorpion to left. ... Made in: Attica, Hellenic republic ... Findspot: Nola, Italy
- ^ Kerényi, C. (1974). "Stories of Orion". The Gods of the Greens. Thames and Hudson. p. 203. ISBN978-0-500-27048-6.
- ^ a b Stockmann & Ythier 2010, p. 179.
- ^ Takeda, A. (2011). "Blumenreiche Handelswege: Ost-westliche Streifzüge auf den Spuren der Fabel Der Skorpion und der Frosch" [Flowery Trade Routes: East-Western forays into the footsteps of the fable The Scorpion and the Frog] (PDF). Deutsche Vierteljahrsschrift für Literaturwissenschaft und Geistesgeschichte (in German language). 85 (one): 124–152. doi:x.1007/BF03374756. S2CID 170169337. Archived (PDF) from the original on five November 2020. Retrieved nine December 2020.
(German language) Die Moral der Fabel besagt: Manche Menschen handeln von Natur aus mörderisch und selbst-mörderisch zugleich. (English) The moral of the fable says: Some people human activity naturally murderous and self-murderous at the same fourth dimension.
- ^ Meyer, Michael (2005). "Diamond in the Rough: Steinbeck'south Multifaceted Pearl". The Steinbeck Review. ii (2 (Fall 2005)): 42–56. JSTOR 41581982.
- ^ Weiss, Allen South. (1996). "Between the sign of the scorpion and the sign of the cross: L'Age d'or". In Kuenzli, Rudolf East. (ed.). Dada and Surrealist Film. MIT Press. pp. 159. ISBN978-0-262-61121-3.
- ^ "Stevie Smith: Bibliography". Poetry Foundation. Archived from the original on one July 2019. Retrieved one July 2019.
- ^ Wallis, J. Doyle (2004). "Operation Scorpio". DVD Talk. Archived from the original on 20 June 2015. Retrieved 19 June 2015.
- ^ "The Scorpion King". Box Office Mojo. Archived from the original on 16 December 2009. Retrieved 4 Jan 2010.
- ^ Provo, Frank (2002). "The Scorpion King: Sword of Osiris Review". GameSpot. Archived from the original on 29 June 2020. Retrieved 24 June 2020.
- ^ a b Iyengar, B. K. S. (1991). Low-cal on Yoga. Thorsons. pp. 386–388. ISBN978-0-00-714516-4. OCLC 51315708.
- ^ Vitruvius, De Architectura, X:ten:one–6.
- ^ "Scorpion". Jane'south Data Group. Archived from the original on 21 Feb 2008. Retrieved 16 November 2020.
- ^ Fletcher, David (2017). British Battle Tanks: British-fabricated Tanks of World War II. Bloomsbury. p. 37. ISBN978-ane-4728-2003-7. Archived from the original on 31 July 2020. Retrieved 27 June 2020.
- ^ Winfield, Rif (2008). British Warships in the Historic period of Sail 1793–1817: Design, Structure, Careers and Fates. Seaforth. p. 291. ISBN978-1-86176-246-vii.
- ^ Parkes, Oscar (1990). British Battleships (reprint of the 1957 ed.). Naval Institute Press. pp. 78–79. ISBN1-55750-075-4.
- ^ "[untitled]". Marine Review. Cleveland, Ohio. fourteen (11): 1. x September 1896.
- ^ "[untitled]". Naval and Armed forces Intelligence. The Times. London. 31 August 1910. p. five. Archived from the original on 31 May 2021. Retrieved 2 December 2020.
- ^ "USS Scorpion (SSN 589) May 27, 1968 – 99 Men Lost". Us Navy. 2007. Archived from the original on 12 March 2008. Retrieved 9 April 2008.
- ^ "The History of Abarth's Logo". Museo del marchio italiano. Archived from the original on 3 July 2020. Retrieved 16 November 2020.
The company logo has been the scorpion since the very start; Carlo Abarth did non just want it as a reference to his zodiac sign, only besides because it was an original and hard to imitate logo. At the beginning the scorpion was gratis from whatsoever profile and featured the typo "Abarth & Co.- Torino". In 1954 a shield was added, as symbol of victory and passion;
- ^ Salvadori, Clement (17 January 2019). "Retrospective: 1974–1977 Montesa Cota 247-T". Passenger Magazine. Archived from the original on 27 June 2020. Retrieved 25 June 2020.
Permanyer persisted, congenital larger engines, and in 1965 showed the 247cc engine (21 horsepower at seven,000 rpm) in a Scorpion motocrosser.
- ^ Betimes; Budig, Kathryn (i Oct 2012). "Kathryn Budig Challenge Pose: Scorpion in Forearm Balance". Yoga Journal. Archived from the original on 27 June 2020. Retrieved 25 June 2020.
Sources
- Polis, Gary (1990). The Biology of Scorpions. Stanford University Press. ISBN978-0-8047-1249-1. OCLC 18991506.
- Stockmann, Roland; Ythier, Eric (2010). Scorpions of the World. N. A. P. Editions. ISBN978-2913688117.
- Stockmann, Roland (2015). "Introduction to Scorpion Biology and Ecology". In Gopalakrishnakone, P.; Possani, 50.; F. Schwartz, Eastward.; Rodríguez de la Vega, R. (eds.). Scorpion Venoms. Springer. pp. 25–59. ISBN978-94-007-6403-3.
External links
- American Museum of Natural History - Scorpion Systematics Research Group
- CDC – Insects and Scorpions – NIOSH Workplace Condom and Health Topic
Source: https://en.wikipedia.org/wiki/Scorpion
0 Response to "Show Me Picture of a Scorpion Biting a Baby's Legs"
Postar um comentário